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CURRENT RESEARCH

Shifting Coyote-Fox Interactions During COVID Lockdowns.

 

In the second chapter of my PhD dissertation, I examined shifts in coyote and red fox space use, temporal activity, and site-level activity to understand how changes in human recreation during lockdown impacted interactions between these urban carnivores.

Readyhough, T. S., Cepek, J. D., Byer, N. W., & Moll, R. J. (2026). A disrupted human shield: the COVID lockdown alters carnivore dynamics in favour of a dominant competitor. Proceedings of the Royal Society B: Biological Sciences, 293(2072), 20252890. DOI: 10.1098/rspb.2025.2890

Carnivore Interactions at a Shared Urban Den Site.

 

I mentored University of New Hampshire undergraduate student Caroline Savage on an independent research project investigating wildlife interactions at a shared urban den site in the Cleveland Metroparks (OH, USA). Caroline observed at least 11 mammal species utilizing the den site and documented interactions between competing carnivores (coyotes and red foxes).

Savage, C. E., Readyhough, T. S., Cepek, J. D., & Moll, R. J. (2026). Carnivore interactions at an urban den used by at least 11 mammal species. Urban Naturalist, (12). https://eaglehill.us/URNAonline2/access-pages/012-Moll-note-accesspage.shtml (open access)​​

Using Drones (Unpiloted Aerial Vehicles) to Estimate Moose Densities.

 

I collaborated with University of New Hampshire M.S. student Lily Hall on her work using UAVs (drones) to estimate moose densities in northern New Hampshire. Lily's work focused on the feasibility of collecting infrared imagery with drones to estimate moose density across a large landscape, and integrated rigorous field surveys with Bayesian modeling approaches to provide moose population densities for managers.

Hall, L. M., F. B. Sullivan, T. S. Readyhough, S. A. Burke, M. W. Palace, H. Jones, A. M. Brehm, and R. J. Moll. (2026) Using drones to estimate moose densities at a landscape scale. Journal of Wildlife Management e70251 DOI: 10.1002/jwmg.70251​​

Urban Deer Fecundity in an Actively Managed Population.

 

In the first chapter of my PhD dissertation, I used Structural Equation Models of 20-years of data from an actively managed deer population to investigate drivers of fecundity in an urban population.

Readyhough, T.S., Cepek, J.D., Shaffer, E.E., Dennis, P.M., Byer, N.W., Hausman, C.E., Montgomery, R.A., Moll, R.J. (2024) Unveiling drivers of fecundity in an urban white-tailed deer population over 20 years of active management. Urban Ecosystems 28, 38. DOI: 10.1007/s11252-024-01633-y

U.S. Terrestrial Mammal Research.

 

The Wildlife Modeling and Monitoring Lab (WMML) at the University of New Hampshire recently published an investigation into the status and trends of U.S. terrestrial mammal research since 1900. Andrew Butler and I are co-first authors of this study. 

Readyhough, T.S., Butler, A.R., Stephen, R.B., Hall, L., Heit, D., Poisson, M.K.P., Moll, R.J. (2024). Status and trends in United States terrestrial mammal research since 1900. Mammal Review. DOI: 10.1111/mam.12367

Large Carnivores in the Anthropocene.

 

The Wildlife Modeling and Monitoring Lab (WMML) at the University of New Hampshire recently contributed a chapter to a forthcoming book "Apex predators in the Anthropocene: How humans shape large carnivore populations" from Oxford University Press. Our chapter focuses on the lethal effects of roads on large carnivores. 

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Poisson, M. K. P., Butler, A. R., Dawson, C. R., Ford, A. T., Readyhough, T. S., Scherger, J. K., & Moll, R. J. (2026). Lethal effects of roads on large carnivores. In R. A. Montgomery & R. J. Moll (Eds.), Apex predators in the Anthropocene: How humans shape large carnivore populations (pp. 103–120). Oxford University Press. DOI: 10.1093/9780191996665.003.0006

Zoo-housed Bull Asian Elephant Social Behaviors.

 

Behavioral Research Associate investigating the introduction of two young bull Asian elephants into the existing bachelor herd at Denver Zoo. This research is ongoing.

Schreier, A. L., Davis, M., Joseph, S., Moresco, A., & Readyhough, T. S. (2025). Bull Asian Elephant Welfare Improves through Social Housing. Animal Behaviour and Welfare Cases, (2025), DOI: abwcases20250027.

Readyhough, T. S., Davis, M., Joseph, S., Moresco, A., & Schreier, A. L. (2024). Intervention by an older, dominant male elephant (Elephas maximus) during an agonistic interaction between adolescents at Denver Zoo. In&Vertebrates. (open access)

Readyhough, T. S., Davis, M., Joseph, S., Moresco, A., & Schreier, A. L. (2023). Age and Social History Impact Social Interactions between Bull Asian Elephants (Elephas maximus) at Denver Zoo. Journal of Zoological and Botanical Gardens, 4(1), 191-213. (open access)

 

Readyhough, T. S., Joseph, S., Davis, M., Moresco, A., & Schreier, A. L. (2022). Impacts of socialization on bull Asian elephant (Elephas maximus) stereotypical behavior. Journal of Zoological and Botanical Gardens, 3(1), 113-130. (open access)

Moresco, A., Prado, N., Davis, M., Schreier, A. L., Readyhough, T. S., Joseph, S., ... & Brown, J. L. (2022). Immunoglobulin A and physiologic correlates of well-being in Asian elephants. Journal of Zoological and Botanical Gardens, 3(4), 677-687. (open access).

Schreier, A. L., Readyhough, T. S., Moresco, A., Davis, M., & Joseph, S. (2021). Social dynamics of a newly integrated bachelor group of Asian elephants (Elephas maximus): Welfare implications. Journal of Applied Animal Welfare Science, 26(2), 229-246.

Thevarajah, S. J., Readyhough, T. S., Davis, M., Moresco, A., Joseph, S., & Schreier, A. L. (2021). Nighttime behavior and the length of social relationships in male Asian elephants. Journal of Applied Animal Welfare Science, 26(3), 447-462.

Zoo Lights Impacts on Great Indian Hornbills.

 

The focus of my MS capstone project was a short research project investigating the impacts of Zoo Lights on the behaviors of a potential breeding pair of Great Indian Hornbills at Denver Zoo under the supervision of Dr. Amy Schreier from Regis University. 

Readyhough, T. S., Joseph, S., Vyas, K., & Schreier, A. L. (2022). The effects of Zoo Lights on animal welfare: A case study of great Indian hornbills at Denver Zoo. Zoo Biology, 41(3), 263-270.

Edge Effects on Mantled Howler Monkeys.

 

I assisted Dr. Amy Schreier and her collaborators on a paper examining edge effects on activity budgets and social cohesion of mantled howler monkeys at La Suerte Biological Station in Costa Rica.

Schreier, A. L., Bolt, L. M., Russell, D. G., Readyhough, T. S., Jacobson, Z. S., Merrigan-Johnson, C., & Coggeshall, E. M. (2021). Mantled howler monkeys (Alouatta palliata) in a Costa Rican forest fragment do not modify activity budgets or spatial cohesion in response to anthropogenic edges. Folia Primatologica, 92(1), 49-57.

Stable Isotope Analysis of Urban Coyote Diets.

 

Undergraduate research with Dr. Seth Newsome, from the University of New Mexico, investigating anthropogenic resource use by urban coyotes in the Chicago area.

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